diff --git a/js/input/Hotkey.ts b/js/input/Hotkey.ts
index c0e704efd..72f99baf2 100644
--- a/js/input/Hotkey.ts
+++ b/js/input/Hotkey.ts
@@ -49,17 +49,19 @@ type SelfOptions = {
   // A set of modifier keys that:
   //
   // 1. Need to be pressed before the main key before this hotkey is considered pressed.
-  // 2. If not a normal (ctrl/alt/meta/shift) modifier key, will also be required to be "off" for other hotkeys to be
-  //    activated when this hotkey is present.
+  // 2. Must NOT be pressed for other hotkeys to be activated when this hotkey is present.
   //
-  // Note that the release of a modifier key may "activate" the hotkey for "fire-on-hold", but not for "fire-on-down".
+  // A Hotkey will also not activate if the standard modifier keys (ctrl/alt/meta/shift) are pressed, unless they
+  // are explicitly included in the modifierKeys array.
+  //
+  // NOTE: This is a generalization of the normal concept of "modifier key"
+  // (https://en.wikipedia.org/wiki/Modifier_key). It is a PhET-specific concept that allows other non-standard
+  // modifier keys to be used as modifiers. The standard modifier keys (ctrl/alt/meta/shift) are automatically handled
+  // by the hotkey system, but this can expand the set of modifier keys that can be used. When a modifier key is added,
+  // pressing it will prevent any other Hotkeys from becoming active. This is how the typical modifier keys behave and
+  // so that is kept consistent for PhET-specific modifier keys.
   //
-  // NOTE: This is a generalization of the normal concept of "modifier key" (https://en.wikipedia.org/wiki/Modifier_key).
-  // It is a PhET-specific concept that allows other non-modifier keys to be used as modifiers.
-  // The standard modifier keys (ctrl/alt/meta/shift) are automatically handled by the hotkey system, but this can
-  // expand the set of modifier keys that can be used.
-  // When a modifier key is added, pressing it will prevent any other Hotkeys from becoming active. This is how the
-  // typical modifier keys behave and so that is kept consistent for PhET-specific modifier keys.
+  // Note that the release of a modifier key may "activate" the hotkey for "fire-on-hold", but not for "fire-on-down".
   modifierKeys?: EnglishKey[];
 
   // A set of modifier keys that can be down and the hotkey will still fire. Essentially ignoring the modifier